The plot of this game is focused on how the history of the Skylands comes alive again in the present. The main villain from the last game, Kaos, finds himself in a toy store. He finds a Portal of Power, and goes back to the Skylands. the first level takes place 10,000 years ago to discover where the Skylanders and the Giants came from. 10,000 years ago, the Skylands were ruled by an evil race of giant robots called Arkeyans, until the Skylanders made a stand and began war with the Arkeyans. Eventually, the Giants, who were the first ever Skylanders, faces the king of the Arkeyans and removes the source of the Arkeyans' power, the Iron Fist of Arkus, but this victory also sent the Giants to Earth, ready for the player to use them.
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